Taxonomic Hierarchy
| Phylum |
Mollusca |
| Class |
Gastropoda
|
| Subclass |
Prosobranchia
|
| Order |
Neogastropoda
|
Superfamily
|
Muricacea |
| Family |
Fasciolariidae Gray, 1853 |
| Subfamily |
Fusininae Wrigley, 1927
|
| Genus |
Granulifusus Kuroda & Habe, 1954 |
Genus Granulifusus Kuroda & Habe, 1954
Type species: Fusus niponicus E.A. Smith, 1879 (Recent, western Pacific) (by monotypy).
The genus Granulifusus is restricted to the upper continental shelves in the Indo-West Pacific and actually contains 22 Recent and 6 fossil species. The taxon was recently discussed by Snyder (2003: 87-89). The shell characteristics cited in the original description of Granulifusus are a granulated surface sculpture and a small round operculum which does not fill the aperture (Kuroda & Habe, 1954: 89). Shuto (1958: 253) cited “… having a few granulated spiral series on the surface and the aperture gently narrowed to the siphonal canal without any constriction.“. Most species are light brown with reddish-brown spiral bands.
Moreover, most Granulifusus species have 1 or 2 indistinct columellar fold(s) visible inside the aperture, often acquired in adult stage.
Periostracum thin and inconspicuous, straw coloured.
Type species of genus Granulifusus Kuroda & Habe, 1954
Different forms of Granulifusus niponicus (E.A. Smith, 1879). G. niponicus is the type species of the genus Granulifusus:
 Granulifusus niponicus (E.A. Smith, 1879), Syntype BMNH 1878.10.16.49, Japan, 22.1 mm |
 Fusus suboblitus Pilsbry, 1904, Holotype ANSP 70941, Japan, 36.5 mm (junior synonym of G. niponicus) |
 Granulifusus niponicus (E.A. Smith, 1879), Japan, Kii, 60.5 mm |
 Fusinus colus (Linnaeus, 1758) |
Recent species
| Taxon |
Range |
| Granulifusus amoenus Hadorn & Fraussen, 2005 |
Vanuatu |
| Granulifusus babae Hadorn & Fraussen, 2005 |
Indonesia, Tanimbar Islands |
| Granulifusus bacciballus Hadorn & Fraussen, 2005 |
New Caledonia, most common in north New Caledonia |
| Granulifusus balbus Hadorn & Fraussen, 2005 |
Southwestern Pacific (south New Caledonia and southern Coral Sea) |
| Granulifusus benjamini Hadorn & Fraussen, 2005 |
Southwestern Pacific (Tonga Islands, Fiji, New Caledonia, Vanuatu, Coral Sea) |
| Granulifusus captivus (E.A. Smith, 1899) |
Andaman Islands |
| Granulifusus consimilis Garrard, 1966 |
Queensland, Australia |
| Granulifusus dondani Snyder, 2003 |
Central Philippine Islands |
| Granulifusus geometricus Hadorn & Fraussen, 2005 |
From Waterwitch Bank and from Tonga Islands to New Caledonia and the Southern Coral Sea |
| Granulifusus hayashii Habe, 1961 |
Japan |
| Granulifusus kiranus Shuto, 1958 |
Western Pacific, from central Honshu in the north to western Australia in the south. Indonesia and New Caledonia |
| Granulifusus libratus (Watson, 1886) |
Southwestern Pacific (Tonga Islands, Fiji, Vanuatu and south New Caledonia) |
| Granulifusus lochi Hadorn & Fraussen, 2005 |
Western Australia, Great Australian Bight |
| Granulifusus monsecourorum Hadorn & Fraussen, 2005 |
Madagascar |
| Granulifusus niponicus (E.A. Smith, 1879) |
Japan (south of central Honshu), Taiwan, Philippine Islands |
| Granulifusus noguchii (Habe & Masuda, 1990) |
Japan, Suruga Bay and southwards, East China Sea, southern Korea |
| Granulifusus poppei Delsaerdt, 1995 |
Somalia |
| Granulifusus pulchellus Hadorn & Chino, 2005 |
Central to southwestern Japan |
| Granulifusus rubrolineatus (Sowerby, II, 1870) |
From South Africa, Agulhas Bank northwards to southern Somalia |
| Granulifusus rufinodis (von Martens, 1901) |
West coast of Sumatra |
| Granulifusus staminatus (Garrard, 1966) |
Southern Pacific (Tonga Islands, Futuna, Fiji, Vanuatu, New Caledonia, southern Coral Sea and Queensland, Australia) and Philippine Islands |
| Granulifusus vermeiji Snyder, 2003 |
Central Philippine Islands |
Fossil species
Taxon
|
Range / Geological Age |
| Granulifusus dualis (Yokoyama, 1928) |
Japan (Early Pliocene) |
| Granulifusus koyuanus Shuto, 1958 |
Japan (Late Miocene) |
| Granulifusus makiyamai (Otuka, 1937) |
Japan (Pleistocene) |
| Granulifusus matsumotoi Shuto, 1958 |
Japan (Early Pliocene) |
| Granulifusus musasiensis (Makiyama, 1922) |
Japan (Pliocene) |
| Granulifusus nakasiensis Hadorn & Fraussen, 2005 |
Fiji, Nakasi Sandstone Beds (Late Pliocene) |
Protoconch
Typical protoconch of Granulifusus is paucispiral, consisting of 1-1 1/2 smooth, glossy whorls, final 1/4 whorl sculptured with some axial riblets, transition to teleoconch abrupt, marked by a varix.
Operculum
The operculum is corneous, thin, translucent, yellowish to brown coloured, round-ovate, smaller than aperture, with fine concentric ridges and with a nucleus located at the lower right hand side.
 Operculum of G. hayashii Habe, 1961 |
 Operculum of G. noguchii (Habe & Masuda, 1990) |
 Operculum of G. rubrolineatus (Sowerby, II, 1870) |
Radula
Radula with a tricuspid central tooth which is rectangular or slightly triangular, lateral teeth curved, with a small number of cusps (4-7), mostly with a denticle at inner side.
 Radula of Granulifusus niponicus (E.A. Smith, 1879) |
 Radula of Granulifusus bacciballus Hadorn & Fraussen, 2005 |
|
Granulifusus
